St Mary and St Peter's Church is in the village of Wilmington, East Sussex, England. It is an active Anglican parish church on the deanery of Lewes and Seaford, the archdeaconry of Lewes and Hastings, and the diocese of Chichester. Founded in the late 11th century to serve villagers in a rural area at the foot of the South Downs, it also functioned as a priory church for the monks from the adjacent Wilmington Priory, to which it was physically connected. The building has "benefited from sympathetic restoration" over the centuries—including a series of works by prominent architects Paley and Austin in the Victorian era, and internal renovation after a fire in the early 21st century. The church is recorded in the National Heritage List for England as a designated Grade I listed building.